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Evidence supporting the paraphyly of Hucho (Salmonidae) based on ribosomal DNA restriction maps. Journal of fish biology. 47:956–961.. 1995.
Phylogenetic analysis of Pacific salmon (genus Oncorhynchus) using nuclear and mitochondrial DNA sequences. Canadian Journal of Fisheries and Aquatic Sciences. 54:1865–1872.. 1997.
Phylogenetic relationships among the Salmoninae based on nuclear and mitochondrial DNA sequences.. Molecular systematics of fishes. Academic Press, London.. 1997.